Greening has been the Member of Parliament (MP) for Putney since the 2005 general election and was appointed Secretary of State for International Development in September 2012. Previously, she was appointed Economic Secretary to the Treasury in May 2010, and became Secretary of State for Transport on 14 October 2011. On 4 September 2012, she was replaced by Patrick McLoughlin at the Department for Transport, and became Secretary of State for International Development.
In February 2013 she was assessed as one of the 100 most powerful women in the United Kingdom by Woman's Hour on BBC Radio 4.
Greening was appointed a vice-chair (with responsibility for youth) of the Conservative Party on 15 December 2005, having earlier that year been appointed a member of the Work and Pensions Committee. In July 2007 following a shadow ministerial reshuffle, she was promoted to be a Junior Shadow Minister for The Treasury.
In January 2009 following a further shadow ministerial reshuffle, Greening was promoted to Shadow Minister for London, within the Communities and Local Government Team with responsibility for Local Government Finance. Within this brief, she focussed on transport and local community benefits.
During the 2009 expenses scandal, Greening was ranked as the 9th best value for money MP in research carried out by the free-market think tank, the Adam Smith Institute, and of all her expenses claims she was the 599th lowest out of the 645 MPs. In March 2010 she was put in charge of co-ordinating the Conservative campaign for the 2010 general election in London. She became Economic Secretary to the Treasury.
In October 2011 she was appointed Secretary of State for Transport and was sworn of the Privy Council.
Greening represents the London constituency of Putney and had always campaigned against a third runway at Heathrow. In the run up to the 2012 Cabinet reshuffle Greening said it would be difficult to serve in a Cabinet which was in favour of a third runway.
In the event Greening was reshuffled on 4 September 2012. The move was attacked by Boris Johnson.
Greening was in a relationship with Mark Clarke, the 2010 Conservative parliamentary candidate in Tooting, who was later expelled from the party for bullying young activists. In June 2016, Greening revealed on Twitter that she was in a "happy same sex relationship". Referring to the pro-remain campaigning group during the EU membership referendum, she added: "I campaigned for Stronger In but sometimes you're better off out!".
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